PM 'is losing command' says Cameron

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Tory leader David Cameron said the fourth ministerial resignation in two days showed the PM had lost command of a Government which was "collapsing before our eyes", while Lib Dem leader Nick Clegg told MPs: "Labour is finished."

But Mr Brown rejected opposition calls for an immediate general election.

He insisted he was focusing on cleaning up the parliamentary expenses system and tackling the recession.

However, he ducked Mr Cameron's challenge to say whether Alistair Darling - tipped for the chop in the reshuffle expected within the next few days - would still be in the Treasury next week.

In a letter to Mr Brown confirming her resignation, Ms Blears said she was proud of her service to the Government over the past eight years and would "work tirelessly to see Labour elected at the next General Election".

But she pointedly offered none of the usual expressions of support for the Prime Minister or his policies.

In response, Mr Brown praised the Salford MP's achievements as a minister and said he hoped she would return to Government in the future.

Mr Cameron repeated his call for an immediate general election in a speech to Conservative activists in Hammersmith, west London.

"People know we have got to cleanse our political system," he said. "We need a general election so the public can be the judge and jury."