Houvenaghel misses out

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Houvenaghel, the fastest qualifier, slowed significantly in the latter part of the women's individual pursuit to finish second to New Zealand's Alison Shanks, who won gold in three minutes 29.807 seconds.

Victoria Pendleton and Chris Newton earlier won bronze medals in the women's 500 metres time-trial and the men's points race, respectively while Britain claimed a fourth successive men's team sprint silver after Jamie Staff, Jason Kenny and Matt Crampton were beaten by France.

Joanna Rowsell could not add a third bronze, finishing fourth in the women's individual pursuit as Lithuania's Vilija Sereikaite clinched third place.

Pendleton, Britain's only defending individual champion, opened her campaign with an unexpected time-trial bronze.

In the first of her four events - she also rides the team sprint with Shanaze Reade, the sprint and the keirin - Olympic sprint champion Pendleton clocked a season's best of 34.102secs in the non-Olympic discipline, with Simona Krupeckaite winning gold in a world record of 33.296secs.

Australia's Anna Meares set a time of 33.796secs to hold first place for much of the event before seeing her world record smashed by the Lithuanian, while Britain's Anna Blyth was 14th in 35.443secs.

Newton won Britain's second bronze of the evening, finishing third in the men's points race.

The 34-year-old, who won Olympic bronze in Beijing, was in contention throughout the 160-lap (40 kilometres) event, which features sprints and points awarded every 10 laps.

However, Australia's Cameron Meyer edged ahead in the later sprints to clinch gold, with Denmark's Daniel Kreutzfeld taking silver.